:D YAY so idr what the th says right now but my current interpretation is that kiophen have at least 4 biosex categories, based on what gamete they make (or don't make). Those are eggs, sperm, and vril, which is an extra gamete that is needed to complete the fertilization. A fertilized egg isn't technically conceived without it. I don't think vril has a real world biological counterpart but the word was a term used for 'life giving elixirs' in the 19th century, and was originally the name of some kind of magical potion/poison from a fictional book called Vril: The Power of the Coming Race (which I have not read, hopefully it's not incredibly racist). It's also short for "virile". Vimales are usually a bit larger and stronger than their other-sex counterparts, and flying is more difficult for them. They're generally better at climbing and digging.

The reason for kimales and 3 gamete sexes is to force kiophen to leave their isolated towns to search for mates, since their small populations can lead to inbreeding quickly. Having 3 parents per organism also mitigates this; recessive traits are statistically less likely to be expressed. Kimales aid in this process in what I called "gene transfer"; they are blessed with equipment that lets them accept gamete materials + half fertilized embryos and bring them to other villages to share.

There are different physical setups a kimale might have, but they usually have some kind of genital that can "Take and Give" gametes. Otherwise, the consistent aspect of kimales is that they don't produce their own gamete. Miles is a kimale, and they have a sort-of hermaphroditic setup, with a womb that connects to both a prehensile ovipositor-tentacle and vagina. Kimales usually have some kind of prehensile tentacle that can worm inside wombs and pump out whatever is inside.

I like the idea of a kiophen being able to chose their genitals, so right now my idea is that every sex starts out null and their Will influences their body type and whatever they got down there. Kiophen have a sort of Orcs From Warhammer 40K thing going on (orcs can alter reality due to their strong combined belief; if they believe/assume something hard enough it becomes true), kiophen just assume they can fly due to gravity being lower in the tree, but it's actually due in part to their telekinetic abilities (they don't really know they have them). So I think it'd be cool if they can change their biology to an extent through their own will.